do you know the co2 concentration when the lights are on and off?
if your co2 bubbling rate gives you 20-30ppm during photoperiod then keeping the same co2 injection rate, will result in co2 overdose.
so your shrimps may be dead due to co2 overdose.
are they any other inhabitants in your tank other than the yamato shrimp?
normally, co2 is injected only during the light on period. a solenoid is used to 'switch on/off' the co2 supply. if you are not using a solenoid, switching on an air pump to outgas excess co2 works as well.
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