I suspect its a lack of certain macro nutrients. However, I wouldn't suggest you dose any fertilizers other than potassium since you have CRS in this tank.
If you can, check your nitrate levels, it may be very low with so many fast growing plants around.
You have to make a decision of whether this tank is for plants or for shrimps, as it is hard to maintain both. I use plants in my shrimp tank only as nitrate and phosphate absorbers which are thrown away regularly.
Mosses grow slowly and are mostly decor or places for shrimplets to hide and feed.
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